Pleased that removing the air filter and pipe has cured the problem. Personaly, I wouldn't try any replacement set-up and would run it as it now is.
Yes, I have already made the change to 428 chain and whilst I did have a lumpier cam in it, I have now returned to a white (standard) cam in the 450, with a less than 10:1 compression and standard diameter valves. So, a "soft" engine.
I have to agree with your final sentence
.I can never put my finger on why Ducati Singles are so great to ride, but possibly it is due to the light weight, torquey, responsive engine, fine steering and that thumping exhaust note in the background.
Good health, Bill